Costa Rica’s subterranean sites: sites with records of cave fauna (red) vs. sites without records of cave fauna (black). The state-protected areas are presented in different colors: state farm (SF), wetland (WL), protected zone (PZ), national park (NP), biological reserve (BR), forest reserve (FR), indigenous reserve (IR), absolute natural reserve (ANR), and wildlife refuge (WR). The maps inserted at the bottom represent two karstic areas in Costa Rica with the most significant clusters of caves A Barra Honda karst area and B Zona Sur karst area.