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Article title
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Abstract
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Keywords
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Introduction
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Methods
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Field collecting
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Morphology
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Material examined
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Taxonomy
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Molecular phylogenetics
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Results
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Molecular phylogenetics
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Taxonomy
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Discussion
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Molecular phylogenetics
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Morphology
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Evidence that Horologion is a trechite
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Evidence that Horologion + Bembidarenini + Trechini form a clade
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Evidence that Horologion + Bembidarenini form a clade
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Evidence that Horologion does not belong in Trechini
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Relationships of Lovriciina
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How aberrant is Horologion?
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Natural history
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Distribution and biogeography
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Conclusions
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Acknowledgements
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References
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Supplementary materials
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