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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Material and methods
    • Study area
    • Environmental features of quartzite caves
    • Faunal survey
    • Descriptions of human uses and impacts
    • Data analyses
  • Results
    • Environmental factors of the caves
    • Faunal composition, abundance, and richness
    • Obligate cave species richness
    • Invertebrate diversity and distribution and their relationships with abiotic factors
    • Trophic resources for the cave invertebrates
    • Description of human uses and impacts
  • Discussion
    • Obligate cave species diversity
    • Threats to the cave fauna
    • Final considerations
  • Acknowledgments
  • References
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