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  • Article title
  • Abstract
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  • Introduction
  • Material and methods
    • Taxon sampling
    • Genetic delimitation
    • Conchological assessment
    • Histology and light microscopy
    • Scanning Electron Microscopy (SEM)
    • X-ray nanotomography
    • Digital imaging
  • Results
    • Genetic delimitation
    • Consequence 1: Zospeum isselianum (L1) neotype designation including shell morphology and genetics
    • Taxonomy
      • Family Carychiidae Jeffreys, 1830 Genus Zospeum Bourguignat, 1856
    • Consequence 2: Histological study of Zospeum sp. (Konečka zijalka) ultrastructure
    • Digestive system
    • Radula
    • Reproductive structures
    • Central nervous system (CNS)
    • Consequence 3: Relegation of Z. alpestre bolei into synonymy of Z. alpestre with differential diagnosis of L3, including context to L2
    • Character analysis of the shell of allied species
    • Consequence 4: Differential diagnosis for Z. kupitzense (L4)
  • Discussion
  • Conclusions
  • Acknowledgements
  • References
  • Supplementary material
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