Detrended Correspondence Analysis (DCA) ordination diagram of benthic species and environmental variables in the cenote X-Batún. Statistical significance based on Monte Carlo test with forward selection p ≤ 0.05. Abbreviations are as follows: temperature (Temp), dissolved oxygen (DO), conductivity (Conductv), total dissolved solids (TDS). G1 assemblage of surface sediments; G2 assemblage of open water sediments; G3 assemblage of cavern; G4 assemblage of cave sediments identified in NMDS. Colors corresponded to that in the NMDS plot.